Wireless sensor network (WSN) has emerged as one of the most promising technologies for the future. This has been enabled by advances in technology and availability of small, inexpensive, and smart sensors resulting in cost effective and easily deployable WSNs. However, researchers must address a variety of challenges to facilitate the widespread deployment of WSN technology in real-world domains. In this survey, we give an overview of wireless sensor networks and their application domains including the challenges that should be addressed in order to push the technology further. Then we review the recent technologies and testbeds for WSNs. Finally, we identify several open research issues that need to be investigated in future. Our survey is different from existing surveys in that we focus on recent developments in wireless sensor network technologies. We review the leading research projects, standards and technologies, and platforms. Moreover, we highlight a recent phenomenon in WSN research that is to explore synergy between sensor networks and other technologies and explain how this can help sensor networks achieve their full potential. This paper intends to help new researchers entering the domain of WSNs by providing a comprehensive survey on recent developments.